1. 2Department of Botany, University of Wisconsin–Madison, 430 Lincoln Drive, Madison, Wisconsin 53706 USA;
2. 3The Morton Arboretum, 4100 Illinois Route 53, Lisle, Illinois 60532‐1293 USA;
3. 4Department of Botany and Laboratories of Analytical Biology, Smithsonian Institution, P.O. Box 37012, NMNH MRC‐166, Washington, D.C. 20013‐7012 USA