1. Department of Microbiology and Infectious Diseases, Kobe University Graduate School of Medicine, Kobe, Japan
2. Graduate School of Pharmaceutical Sciences and Graduate School of Dentistry, Osaka University, Osaka, Japan
3. Division of Mucosal Immunology and International Research and Development Center for Mucosal Vaccines, The Institute of Medical Sciences, The University of Tokyo, Tokyo, Japan
4. Laboratory of Vaccine Materials, National Institute of Biomedical Innovation, Osaka, Japan