1. Woods Hole Research Center, 149 Woods Hole Road, Falmouth, Massachusetts 02540, U.S.A
2. Center for Biodiversity and Conservation, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024, U.S.A
3. Institute of Arctic Biology, Department of Biology and Wildlife, University of Alaska Fairbanks, P.O. Box 80425, Fairbanks, Alaska 99708, U.S.A