1. Department of Biology, College of Science, Bahir Dar University, P.O. Box 79, Bahir Dar, Ethiopia
2. FAO-Sub Regional Office for Eastern Africa, P.O. Box 5536, Addis Ababa, Ethiopia
3. Fisheries and Aquatic Science Stream, Faculty of Life Sciences, Addis Ababa University, P.O. Box 1176, Addis Ababa, Ethiopia