1. Atmosphere and Ocean Research Institute, The University of Tokyo, Kahiwa, Chiba 277-8564, Japan (e-mail: , phone: , fax: ).
2. School of Marine Biosciences, Kitasato University, Sagamihara, Kanagawa 252-0373, Japan.
3. Department of Biology, Tokyo Medical University, Shinjuku, Tokyo 160-8402, Japan.
4. College of Fisheries-Freshwater Aquaculture Center, Central Luzon State University, Science City of Munoz, Nueva Ecija, Philippines.
5. Institute of Fisheries Science, College of Life Science, National Taiwan University, Taipei, Taiwan.