1. Université Clermont Auvergne, CNRS, GEOLAB, F-63000 Clermont-Ferrand, France.
2. Délégation à la Recherche, Government of French Polynesia, B. P. 20981, 98713 Papeete, Tahiti, French Polynesia.
3. Université Clermont Auvergne, PIAF UMR 547, F-63000 Clermont-Ferrand, France.
4. School of Life Sciences, University of Hawai‘i at Manoa, Honolulu, 3190 Maile Way, HI 96822, USA.
5. Department of Geography, Université Clermont Auvergne, F-63000 Clermont-Ferrand, France,
6. University of French Polynesia, Faa'a, Tahiti, French Polynesia.