1. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, IA, USA.
2. The Ecology of Bird Loss Project, Mangilao, GU, USA.
3. Wyoming Cooperative Fish and Wildlife Research Unit, Department of Zoology and Physiology, University of Wyoming, Laramie, WY, USA.
4. Western Pacific Tropical Research Center, University of Guam, Mangilao, GU, USA.