1. Biodiversity Institute and Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045, USA.
2. U.S. Geological Survey, 4165 Spruance Road, Suite 200, San Diego, CA 92101, USA.
3. Ecological Solutions Solomon Islands, Gizo Island, Western Province, Solomon Islands.
4. Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87106, USA.