1. Zoology Department, University of Hawai'i at Mānoa, 2538 McCarthy Mall, Honolulu, Hawai'i 96822.
2. Université Pierre et Marie Curie-Paris 6, CNRSUMR 7144 AD2M Station Biologique, BP 74 29682 Roscoff Cedex, France.
3. Water Resources Research Center, University of Hawai'i at Mānoa, Honolulu, Hawai'i 96822.