1. National Park Service, Pacific Island Inventory and Monitoring Network, P.O. Box 52, Hawai‘i National Park, HI 96718, USA.
2. U.S. Geological Survey, Pacific Island Ecosystems Research Center, P.O. Box 44, Hawai‘i National Park, HI 96718, USA.
3. Department of Biology, University of Hawai‘i at Hilo, 200 W. Kawili St., Hilo, HI 96720, USA.