1. Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, 16-2 Onogawa, Tsukuba, Ibaraki 305-8506, Japan.
2. Institute of Boninology, Nishi-machi, Chichijima, Ogasawara, Tokyo 100-2101, Japan.
3. Graduate School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, 606-8502, Japan.