1. Department of Biology, University of Hawai‘i at Mänoa, Honolulu, Hawai‘i.
2. Pacific Biosciences Research Center, School of Ocean and Earth Science and Technology, University of Hawai‘i at Mänoa, Honolulu, Hawai‘i.
3. Current address: College of Natural and Computational Sciences, Hawai‘i Pacific University, 45-045 Kamehameha Highway, Käne‘ohe, Hawai‘i 96744-5297.