1. Ministry of Agriculture, Tropical Weed Research Unit, Plant Protection Section, Koronivia Research Station, P.O Box 77, Nausori, Republic of Fiji.
2. University of Queensland, Tropical and Subtropical Weed Research Unit, The School of Agriculture and Food Sciences, St. Lucia, Queensland, 4072, Australia.
3. Department of Agriculture, Fisheries, and Forestry, Biosecurity Queensland, Ecosciences Precinct, GPO Box 267, Brisbane, Queensland 4001, Australia.