1. U.S. Geological Survey, Fort Collins Science Center, Department of Biology, MSC03 2020, University of New Mexico, Albuquerque, New Mexico 87131-0001 (e-mail: ).
2. Washington Department of Fish and Wildlife, 600 Capitol Way North, Olympia, Washington 98501-1091.
3. U.S. Geological Survey, Fort Collins Science Center, 2150 Centre Avenue, Building C, Fort Collins, Colorado 80526.