1. United States Department of Agriculture, Animal and Plant Health Inspection Service, Plant Protection and Quarantine, The Ohio State University, Columbus, Ohio 43212 USA
2. United States Department of Agriculture, Animal and Plant Health Inspection Service, Plant Protection and Quarantine, National Museum of Natural History, Smithsonian Institution,Washington, DC 20560, USA
3. Centro Zamorano de Biodiversidad, Escuela Agrícola Panamericana, El Zamorano, Honduras