1. a State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China
2. b Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China
3. c Institution of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences, Guangzhou, China
4. d University of Chinese Academy of Sciences, Beijing, China
5. e Guilin University of Electronic Technology, Beihai, China