1. State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China
2. State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, and Institution of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences, Guangzhou, China, and School of Marine Sciences, University of Maine, Orono, Maine
3. Application Laboratory, Japan Agency for Marine-Earth Science and Technology, Yokohama, Japan