1. Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Japan
2. Department of Natural History Sciences, Graduate School of Science, Hokkaido University, Sapporo, Japan
3. Faculty of Environmental Earth Science, Hokkaido University, Sapporo, and Research Institute for Global Change, Japan Agency for Marine-Earth Science and Technology, Yokohama, Japan
4. Research Institute for Global Change, Japan Agency for Marine-Earth Science and Technology, Yokohama, Japan