1. ECRC, University College London, 26 Bedford Way, London, WC1H 0AP, UK
2. Department of Geography, Lancaster University, Lancaster, LA1 4YB, UK
3. Dept. of Geography, University of Edinburgh, Drummond Street, Edinburgh, EH8 9XP, UK
4. OAFD Freshwater Fisheries Laboratory, Faskally, Pitlochry, PH16 5LB, UK