1. Department of Ichthyology, The Academy of Natural Sciences of Drexel University, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103–1195, USA Email:
2. Department of Ichthyology, The Academy of Natural Sciences of Drexel University, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103–1195, USA
3. Laboratório de Ictiologia de Altamira, Universidade Federal do Pará, Rua Coronel José Porfírio, 2515, Altamira, Pará, 68372-040, Brazil Email: