1. (HRL, XYK) CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Guangzhou 510301, China, e-mail: luohairong14@mails.ucas.ac.cn; xykong@scsio.ac.cn
2. (TAM) National Systematics Laboratory, NOAA/NMFS/OS&T, Post Office Box 37012, Smithsonian Institution, NHB, MRC-153, Washington, DC 20013-7012, U.S.A., e-mail: munroet@si.edu