1. *Department of Molecular Biology and Biochemistry, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka 565-0871, Japan;
2. Department of Immunobiology, School of Pharmacy, Iwate Medical University, Yahaba, Shiwa-gun, Iwate 028-3694, Japan;
3. Department of Health Chemistry, Graduate School of Pharmaceutical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan;
4. The Center for Electron Microscopy and Bio-Imaging Research, Iwate Medical University, Morioka, Iwate 020-8505, Japan; and
5. Department of Physiology, School of Medicine, Tokyo Women's Medical University, Shinjuku-ku, Tokyo 162-8666, Japan