1. Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Hongo, Tokyo 113-0033, Japan
2. Genetic Dynamics Research Unit-Laboratory, The Institute of Physical and Chemical Research (RIKEN), Wako-shi, Saitama 351-0198, Japan
3. Cellular & Molecular Biology Laboratory, The Institute of Physical and Chemical Research (RIKEN)/CREST of the JST, Wako-shi, Saitama 351-0198, Japan