1. School of Biochemistry and Genetics, The Medical School, University of Newcastle, Newcastle-upon-Tyne NE2 4HH, United Kingdom;
2. Cancer Research UK Cell Regulation Group, Paterson Institute for Cancer Research, Christie Hospital NHS Trust, Manchester M20 4BX, United Kingdom; and
3. Division of Yeast Genetics, National Institute for Medical Research, London NW7 1AA, United Kingdom