1. Hanwoo Research Institute, National Institute Animal Science, Rural Development Administration, Pyeongchang 25340, Korea
2. Animal Genetic Resources Research Center, National Institute Animal Science, Rural Development Administration, Hamyang 50000, Korea
3. Poltury Research Institute, National Institute Animal Science, Rural Development Administration, Pyeongchang 25342, Korea
4. Department of Animal Resources Science, Kongju National Universuty, Yesan 32439, Korea
5. Animal Biotechnology Division, National Institute Animal Science, Rural Development Administration, Wanju 55365, Korea