1. Cooperative Institute for Marine and Atmospheric Research, School of Ocean and Earth Science and Technology, University of Hawai‘i at Mānoa 1 , Honolulu, Hawaii 96822, USA
2. Saltwater Inc 2 ., Anchorage, Alaska 99501, USA
3. Pacific Islands Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration 3 , Honolulu, Hawaii 96818, USA
4. Centre for Research into Ecological and Environmental Modelling, The Observatory, University of St Andrews 4 , St Andrews, KY16 9LZ, Scotland
5. Departamento de Biologia Animal, Centro de Estatística e Aplicações, Faculdade de Ciências da Universidade de Lisboa 5 , Portugal