1. Biology Department, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts 02543, USA
2. Department of Natural Sciences, University of South Carolina Beaufort, Bluffton, South Carolina 29909, USA
3. Raytheon BBN, Cambridge, Massachusetts, 02138, USA
4. Key Laboratory of Underwater Acoustic Communication and Marine Information Technology of the Ministry of Education, College of Ocean and Earth Sciences, Xiamen University, Xiamen 361005, China