1. Zhejiang Provincial Key Laboratory for Genetic Improvement and Quality Control of Medicinal Plants, Hangzhou Normal University, Hangzhou 311121, China
2. CAS Key Laboratory for Plant Biodiversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
3. Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Science, Menglun 666303, Yunnan, China