1. Department of Forestry and Environmental Resources, North Carolina State University, Campus Box 8008, Raleigh, NC 27695-8008, U.S.A.
2. Department of Plant and Microbial Biology, North Carolina State University, Campus Box 7612, Raleigh, NC 27695-7612, U.S.A.
3. Institute for Biodiversity Science and Sustainability, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U.S.A.
4. Department of Biology, Appalachian State University, 572 Rivers Street, Boone, NC 28608, U.S.A.
5. University of North Carolina Herbarium, CB #3280 Coker Hall, 120 South Road, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, U.S.A.