1. National Institute for Amazonian Research, Department of Biodiversity, Av. André Araújo, 2936, Petrópolis, CEP 69060-001, Manaus, Amazonas, Brazil
2. University of Brasília, Botany Department, Campus Universitário Darcy Ribeiro, Brasília, Distrito Federal, Brazil
3. Graduate Program in Biotechnology and Natural Resources, School of Health Sciences, State University of Amazonas, CEP 69065-001, Manaus, Amazonas, Brazil
4. University of Florida, Department of Biology, Gainesville, FL 32611-8525, U.S.A.