1. School of Life Sciences, East China Normal University, 500 Dongchuan Road, Shanghai 200241, China;
2. Forest Research Centre, Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia;
3. Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia