1. Federal University of Mato Grosso do Sul, Institute of Biosciences, Laboratory of Ecology and Evolutionary Biology, Av. Costa e Silva, s/n, 79070-900, Campo Grande, MS, Brazil
2. Federal University of Mato Grosso do Sul, CPAQ, Av. Oscar Trindade de Barros 740, 79200-000, Aquidauana, MS, Brazil
3. Federal University of Mato Grosso do Sul, Institute of Biosciences, Laboratory of Botanic / Lichenology, Av. Costa e Silva, s/n, 79070-900, Campo Grande, MS, Brazil