1. Laboratory of Plant Ecology and Geoecology, Institute of Marine Geology and Geophysics, Far Eastern Branch of the Russian Academy of Sciences, Nauki, 1B, Yuzhno-Sakhalinsk, 693022, Russia E-mail:
2. Department of Natural History, Bergen University Museum, Box 7800, Allégt.41, N'5012 Bergen, Norway E-mail: