1. Molecular Biology Institute and Department of Molecular, Cell, and Developmental Biology, University of California at Los Angeles, Los Angeles, California 90095
2. Faculty of Bioscience and Biotechnology, Tokyo Institute of Technology and Frontier Collaborative Research Center, Yokohama, 226-8503, Japan
3. Infectious and Inflammatory Disease Center, Burnham Institute for Medical Research, 10901 North Torrey Pines Road, La Jolla, California 92037