1. Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, University of Tokyo, Tokyo, Japan
2. Graduate School of Medicine, University of Tokyo, Tokyo, Japan
3. Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan
4. Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan
5. Department of Molecular Virology, Tokyo Medical and Dental University, Tokyo, Japan
6. CREST, Japan Science and Technology Agency, Saitama, Japan