1. Department of Sustainable Agriculture Science, Rothamsted Research, North Wyke, Devon, United Kingdom
2. Department of Computational and Analytical Sciences, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom
3. Department of Sustainable Agriculture Science, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom
4. Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, USA