1. BGI-ShenzhenShenzhen 518083, China
2. State Key Laboratory of Pathogen and BiosecurityBeijing Institute of Microbiology and EpidemiologyBeijing 100071, China
3. Institute of Medical Microbiology, Virology and HygieneUniversity Medical Centre Hamburg-EppendorfMartinistrasse 5220246 Hamburg, Germany
4. Liaoning UniversityShenyang 110036, China