1. State Key Laboratory of Biocontrol, Southern Marine Sciences and Engineering Guangdong Laboratory (Zhuhai), School of Marine Sciences, Sun Yat-sen University, Guangzhou, China
2. School of Ecology, Sun Yat-sen University, Guangzhou, China
3. School of Life Sciences, Sun Yat-sen University, Guangzhou, China
4. State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou, China