1. Department of Environmental Science, University of Arizona, Tucson, Arizona, USA
2. Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA
3. Cooperative Institute for Research in Environmental Sciences, University of Colorado, Boulder, Colorado, USA
4. Department of Microbiology, The Ohio State University, Columbus, Ohio, USA
5. Department of Applied Ecology, North Carolina State University, Raleigh, North Carolina, USA