1. Program in Cellular and Molecular Biology, University of Wisconsin—Madison, Madison, Wisconsin, USA
2. Department of Biostatistics and Medical Informatics, University of Wisconsin—Madison, Madison, Wisconsin, USA
3. Department of Statistics, University of Wisconsin—Madison, Madison, Wisconsin, USA
4. Laboratory of Genetics, University of Wisconsin—Madison, Madison, Wisconsin, USA
5. Genome Center of Wisconsin, University of Wisconsin—Madison, Madison, Wisconsin, USA