1. Department of Plant Pathology, Nanjing Agricultural University, Nanjing, P. R. China
2. Yunnan Academy of Tobacco Agricultural Sciences, Key Laboratory of Tobacco Biotechnological Breeding, National Tobacco Genetic Engineering Research Center, Kunming, China
3. Yunnan Provincial Key Laboratory of Agri-Biotechnology, Institute of Biotechnology and Genetic Resources, Yunnan Academy of Agricultural Sciences, Kunming, Yunnan, P. R. China
4. Laboratory of Virology, Department of Plant Sciences, Wageningen University & Research, Wageningen, The Netherlands