1. WHO Collaborating Centre for Reference and Research on Influenza, 10 Wreckyn St., North Melbourne, Victoria 3051, Australia
2. Monash University, School of Applied Sciences, Churchill, Victoria 3842, Australia
3. Vaccine and Immunisation Research Group, University of Melbourne, Melbourne School of Population Health and Murdoch Childrens Research Institute, Royal Childrens Hospital, Melbourne, Victoria 3010, Australia
4. University of Oxford, Department of Zoology, Institute for Emergent Infections of Humans, James Martin 21st Century School, Oxford OX1 3PS, United Kingdom