1. College of Food Science, Sichuan Agricultural University, Ya’an, Sichuan, China
2. Department of Microbiology, Faculty of Agriculture and Forestry, University of Helsinki, Helsinki, Finland
3. Laboratory of Gene Technology, Division of Animal and Human Health Engineering, Department of Biosystems, Katholieke Universiteit Leuven, Leuven, Belgium
4. Organismal and Evolutionary Research Programme, Faculty of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland