1. Biotechnology Research Center, Toyama Prefectural University, Toyama 939-0398, Japan
2. Kyowa Hakko Bio Co. Ltd., Chiyoda-ku, Tokyo 100-8185, Japan
3. Division of Agriculture and Agricultural Life Science, The University of Tokyo, Bunkyo-ku, Tokyo 113-8657, Japan
4. Faculty of Applied Bioscience, Tokyo University of Agriculture, Setagaya-ku, Tokyo 156-8502, Japan