1. Earth and Environmental Science, New Mexico Institute of Mining and Technology, Socorro, New Mexico, USA
2. National Cave and Karst Research Institute, Carlsbad, New Mexico, USA
3. Department of Geosciences, The Pennsylvania State University, University Park, Pennsylvania, USA
4. Department of Biology, University of New Mexico, Albuquerque, New Mexico, USA
5. Biological Sciences Academic Division, Autonomous University Juárez de Tabasco, Villahermosa Centro, Tabasco, Mexico