1. Marine College, Shandong University, Weihai, People’s Republic of China
2. Institute for Environmental Genomics, University of Oklahoma, Norman, Oklahoma, USA
3. State Key Laboratory of Microbial Technology, Institute of Microbial Technology, Shandong University, Qingdao, China
4. State Key Joint Laboratory of Environment Simulation and Pollution Control, School of Environment, Tsinghua University, Beijing, China