1. Department of Physiology and Cell Biology, Graduate School of Medicine, Kobe University, 7-5-1 Kusunoki-cho, Chuo-ku, Kobe 650-0017, Japan
2. Okazaki Institute for Integrative Biosciences, National Institutes of Natural Sciences
3. Department of Basic Biology, Graduate School of Advanced Studies (SOKENDAI), 5-1 Higashiyama, Myodaiji, Okazaki, Aichi 444-8787, Japan
4. Department of Molecular Biology and Biochemistry, Graduate School of Medicine, Faculty of Medicine, Osaka University, 2-2 Yamadaoka, Suita, Osaka 565-0871, Japan