1. TBI, Université de Toulouse, CNRS, INRAE, INSA, Toulouse, France
2. Institute of Biological Environmental and Rural Sciences, Aberystwyth University, Aberystwyth, United Kingdom
3. CNRS UMR 7257, Aix-Marseille University, Marseille, France
4. INRAE, USC 1408 AFMB, Marseille, France
5. Department of Biological Sciences, King Abdulaziz University, Jeddah, Saudi Arabia