1. Department of Aquatic Animal Medicine, Research Center for Marine Biology, College of Fisheries, Huazhong Agricultural University, Wuhan, China
2. School of Life Sciences, Beijing Normal University, Beijing, China
3. National Institute of Biological Sciences, Beijing, China
4. State Key Laboratory of Marine Resource Utilization in South China Sea, Key Laboratory of Tropical Biological Resources of Ministry of Education, Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresources, College of Marine Science, Hainan University, Haikou, China