1. Department of Microbiology, Meiji Pharmaceutical University, Noshio, Kiyose, Tokyo, Japan
2. Department of Microbial Science and Host Defense, Meiji Pharmaceutical University, Noshio, Kiyose, Tokyo, Japan
3. Japan Collection of Microorganisms, RIKEN BioResource Center, Koyadai, Tsukuba, Ibaraki, Japan
4. Division of Genomic Technologies, RIKEN Centre for Life Science Technologies, Suehiro-cho, Tsurumi-ku, Yokohama, Kanagawa, Japan